These trucks have two large wheels at front and tank treads at back. These allow them to move across difficult terrain. These trucks have a single exhaust pipe. Inside the truck's cargo storage area are rotating shelves for storaging WALL•Es. The truck is used for moving WALL•Es from place to another after they have cleaned one area. It is also probably used to store them at night when they are inactive.
The last active WALL•E still uses one of these trucks as his home after 700 years. Because the other WALL•Es are no longer active and not in the truck, there is a lot of free space. WALL•E has placed there all the interesting items he has found from the mountains of trash. He also stores important spare parts for himself.
The truck gets damaged twice during the movie by EVE. When WALL•E tries to teach EVE to dance, she accidentally knocks WALL•E at a wall of the truck, damaging both WALL•E and the truck. At the second time she blasts a hole on the ceiling of the truck after she have repaired WALL•E and he needed light for recharge.